- PhD, Biology (Dissertation advisor: Dr. Raleigh J. Robertson), Queen’s University, Kingston, Ontario, Canada, 1988-1992
- MS, Biology (Thesis advisor: Dr. Robert B. Payne), University of Michigan, Ann Arbor, Michigan, 1985-1988
- BS, Biology, University of Michigan, Ann Arbor, Michigan, 1980-1983
Evolution, ecology, animal behavior, ornithology and systematics. Using field studies and genetic techniques (e.g. DNA sequencing, microsatellites, and DNA fingerprinting) to study evolutionary questions. In particular, how ecological and phylogenetic factors influence the evolution and maintenance of behavioral patterns (e.g. parental investment, mating system, mate choice) and life-history characteristics.