{"id":3940,"date":"2016-11-30T08:53:34","date_gmt":"2016-11-30T14:53:34","guid":{"rendered":"https:\/\/uwm.edu\/field-station\/?p=3940"},"modified":"2017-03-11T10:41:59","modified_gmt":"2017-03-11T16:41:59","slug":"brown-lacewing","status":"publish","type":"post","link":"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/","title":{"rendered":"Brown Lacewing (Family Hemerobiidae)"},"content":{"rendered":"<p>Hey, BugFans,<\/p>\n<p>One of the last bugs to grace the BugLady\u2019s porch light in autumn is the brown lacewing (she sees its jewel-like cousin, the green lacewing, in mid-to-late summer). Because of this year\u2019s long, mild fall, brown lacewings straggled on into early November.  <\/p>\n<h3>Brown Lacewings<\/h3>\n<p>Brown lacewings belong to the order Neuroptera (&#8220;nerve-winged\u201d). The order seems to be under constant review by taxonomists, and its inhabitants have been shuffled and re-dealt into suborders under the superorder Neuropterida, which embraces lacewings as well as oddballs like <a href=\"http:\/\/bugguide.net\/node\/view\/324938\/bgpage\">mantidflies<\/a>  (yes, these occur in Wisconsin), <a href=\"http:\/\/bugguide.net\/node\/view\/432609\/bgpage\">spongillaflies<\/a>  (so do these), <a href=\"http:\/\/bugguide.net\/node\/view\/288349\/bgpage\">antlions<\/a>  (and these), and <a href=\"http:\/\/bugguide.net\/node\/view\/824399\/bgpage\">owlflies<\/a>. Snakeflies and the more familiar alderflies, fishflies, and dobsonflies (hellgrammites) have been dealt out and given their own orders but are sometimes included in discussions about the Neuropterida. Neuropterans are generally soft-bodied, weak-flying insects (they\u2019ve been likened to drab, awkward damselflies) that are active in the dusk\/dark and that have complete metamorphosis (egg-larva-pupa-adult). Brown lacewings are in the family Hemerobiidae, with about 500 species worldwide, 60 in the U.S. <\/p>\n<p>[metaslider id=3946]<\/p>\n<p>Adult brown lacewings have four-wings and are a half-inch-long(ish), with light brown wings, often patterned, and they are less conspicuous&mdash;and hairier&mdash;than green lacewings (family Chrysopidae). They have chewing mouthparts and conspicuous eyes, and they somewhat resemble caddisflies. Despite what some books say, the stiletto-shaped larvae of brown lacewings do not (cannot, because they lack dorsal <em>setae<\/em> and <em>tubercles<\/em> to stick things to), disguise themselves by fastening the dry shells of their prey to their backs (green lacewing larvae do and are sometimes called \u201ctrash bugs\u201d for their troubles). Larvae use their long abdomen as an aid in climbing, and they have <a href=\"http:\/\/bugguide.net\/node\/view\/471809\">pincer-like mouthparts designed for piercing their prey and then sucking out the juices<\/a>.  <\/p>\n<p>[metaslider id=3964]<\/p>\n<p>Brown lacewing adults and larvae richly deserve their nicknames of \u201caphid lions\u201d and \u201caphid wolves,\u201d but they also prey on small critters like mealybugs, white flies, spider mites, scales, and on insect eggs (including those of their confreres). Despite their good camouflage, brown lacewings are eaten by birds, ladybugs (fellow aphid-eaters), dragonflies, and spiders.  <\/p>\n<p>Because their list of food items includes some agricultural pests, and because the adults are relatively long-lived (several months) and the females relatively fecund, they have been deployed to act as biological controls (you can buy them). In a 1923 \u201cbiography\u201d of a brown lacewing named <em>Micromus posticus<\/em> (much of the brown lacewing literature is pretty dusty), Ohio Agricultural Station researcher Clifford R. Cutright noted that a brown lacewing may eat 41 aphids during its larval stage and an adult female may lay more than 450 eggs.  <\/p>\n<p>According to &#8220;The Life Histories and Stages of Some Hemerobiids and Allied Species (Neuroptera)\u201d by Roger Smith, in the <em>Annals of the Entomological Society of America<\/em>, 1923, <\/p>\n<blockquote><p>\nA very noticeable feature of females ready to oviposit is the large and much-distended abdomen (the BugLady wonders if the plain-winged brown lacewing pictured here might be a female). The abdomen may assume a salmon or a light amber coloration between the sclerites because of the eggs within. They walk about excitedly, stop suddenly, and bend the abdomen forward, arching it in the middle. They walk about excitedly, stop suddenly, and bend the adomen forward, arching it in the middle. The egg&#8230; is deposited flat on the substratum to which it adheres. The time required for this performance is but a few seconds.\n<\/p><\/blockquote>\n<p>Green lacewing eggs are set like tiny knobs on the end of stiff stalks, but brown lacewing eggs are laid directly on leaf surfaces, buds, bark, etc. near aphid populations. The larvae are active and fleet of foot in the first instar but may become more sedentary in the second two instars. One reference said that larvae whip their heads from side to side, a behavior that the BugLady saw in a larva that ran through a herd of aphids, brandishing one of their fallen comrades. They often hunt on milkweed leaves, but they prefer aphids that feed on plant juices from milkweed species whose <em>cardenolide<\/em> (toxin) levels are low. Mature larvae spin loose, double-walled cocoons in sheltered areas; there may be several generations per year, and the <a href=\"http:\/\/bugguide.net\/node\/view\/675253\">final brood may overwinter as a larva or a pupa<\/a> (or as an adult in Southern climes).  <\/p>\n<p><a href=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/11\/grn-lacewing-5-425.jpg\"><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/11\/grn-lacewing-5-425.jpg\" alt=\"grn-lacewing-5-425\" width=\"425\" height=\"700\" class=\"aligncenter size-full wp-image-3954\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/11\/grn-lacewing-5-425.jpg 425w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/11\/grn-lacewing-5-425-182x300.jpg 182w\" sizes=\"auto, (max-width: 425px) 100vw, 425px\" \/><\/a><\/p>\n<p>The BugLady thinks that a few of her lacewings with streaky wings might be <em>Micromus posticus<\/em>, a species that\u2019s found across North America but seems more common in the Great Lakes area. This species was featured in an article suggesting that an increasing number of sightings of the species in Quebec could be part of a northern migration made possible by global climate change. The authors pointed out that the status of fall-occurring species can be hard to determine because systematic insect collecting drops off after September.  <\/p>\n<p>Interesting brown lacewing factoids:<\/p>\n<ul>\n<li>Adults feign death when alarmed, tucking their \u201cchins\u201d down and their antennae between their stiffened legs.<\/li>\n<li>Their intestines are plugged and they don\u2019t poop until they are adults (not uncommon in this group). A silk-spinning gland occupies the larval posterior.<\/li>\n<li>Larvae \u201cglue\u201d themselves to the substrate by the tip of the abdomen while molting so they won\u2019t fall during ecdysis.<\/li>\n<li>Using their middle and rear pairs of legs, brown lacewings can jump into the air like a grasshopper on their way to becoming airborne. They leap first, and then they flap. Because they use four legs, they can launch from surfaces that might \u201cgive\u201d beneath them, because their weight is spread out. <\/li>\n<\/ul>\n<p>&nbsp;<br \/>\n<em>The BugLady<\/em><\/p>\n","protected":false},"excerpt":{"rendered":"<p>Adult <strong>Brown Lacewings<\/strong> have four-wings and are a half-inch-long(ish), with light brown wings, often patterned, and they are less conspicuous&mdash;and hairier&mdash;than green lacewings. They have chewing mouthparts and conspicuous eyes. Brown lacewing adults and larvae richly deserve their nicknames of \u201caphid lions\u201d and \u201caphid wolves,\u201d but they also prey on small critters like mealybugs, white flies, spider mites, scales, and on insect eggs.<\/p>\n","protected":false},"author":28,"featured_media":0,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":{"_acf_changed":false,"footnotes":"","uwm_wg_additional_authors":[]},"categories":[8],"tags":[267],"class_list":["post-3940","post","type-post","status-publish","format-standard","hentry","category-bug-of-the-week","tag-lacewings"],"yoast_head":"<!-- This site is optimized with the Yoast SEO Premium plugin v27.2 (Yoast SEO v27.2) - https:\/\/yoast.com\/product\/yoast-seo-premium-wordpress\/ -->\n<title>Field Station<\/title>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Brown Lacewing (Family Hemerobiidae)\" \/>\n<meta property=\"og:description\" content=\"Adult Brown Lacewings have four-wings and are a half-inch-long(ish), with light brown wings, often patterned, and they are less conspicuous&mdash;and hairier&mdash;than green lacewings. They have chewing mouthparts and conspicuous eyes. Brown lacewing adults and larvae richly deserve their nicknames of \u201caphid lions\u201d and \u201caphid wolves,\u201d but they also prey on small critters like mealybugs, white flies, spider mites, scales, and on insect eggs.\" \/>\n<meta property=\"og:url\" content=\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/\" \/>\n<meta property=\"og:site_name\" content=\"Field Station\" \/>\n<meta property=\"article:published_time\" content=\"2016-11-30T14:53:34+00:00\" \/>\n<meta property=\"article:modified_time\" content=\"2017-03-11T16:41:59+00:00\" \/>\n<meta property=\"og:image\" content=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/11\/grn-lacewing-5-425.jpg\" \/>\n<meta name=\"twitter:card\" content=\"summary_large_image\" \/>\n<meta name=\"twitter:label1\" content=\"Written by\" \/>\n\t<meta name=\"twitter:data1\" content=\"\" \/>\n\t<meta name=\"twitter:label2\" content=\"Est. reading time\" \/>\n\t<meta name=\"twitter:data2\" content=\"5 minutes\" \/>\n<script type=\"application\/ld+json\" class=\"yoast-schema-graph\">{\"@context\":\"https:\/\/schema.org\",\"@graph\":[{\"@type\":\"Article\",\"@id\":\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/#article\",\"isPartOf\":{\"@id\":\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/\"},\"author\":{\"name\":\"\",\"@id\":\"\"},\"headline\":\"Brown Lacewing (Family Hemerobiidae)\",\"datePublished\":\"2016-11-30T14:53:34+00:00\",\"dateModified\":\"2017-03-11T16:41:59+00:00\",\"mainEntityOfPage\":{\"@id\":\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/\"},\"wordCount\":921,\"image\":{\"@id\":\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/#primaryimage\"},\"thumbnailUrl\":\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/11\/grn-lacewing-5-425.jpg\",\"keywords\":[\"Lacewings\"],\"articleSection\":[\"Bug of the Week\"],\"inLanguage\":\"en-US\"},{\"@type\":\"WebPage\",\"@id\":\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/\",\"url\":\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/brown-lacewing\/\",\"name\":\"Brown Lacewing (Family Hemerobiidae) - 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